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Anacamptis papilionacea ssp papilionacea
 

The A. papilionacea group is a small but highly distinct collection of orchids that are not easily mistaken for any other. It consists of just two species, A. papilionacea itself, commonly known as the Pink Butterfly Orchid and the Fan Lipped Orchid, A. collina.

A. papilionacea is a polymorphic species with a large and varied distribution, factors that unsurprisingly give rise to a wide range of natural and regional variations. First described (as an Orchis) by Linnaeus as long ago as 1759, the taxon has ever since been intensely studied, leading to the recognition of many forms and the creation of a list of synonyms far too extensive to detail in these pages (67 in 2013). A genetic study in 1993 comparing sub-species grandiflora, papilionacea and heroica (now aegaea) determined that the minimal genetic difference's discovered, were consistent with separate geographic populations within a single cohesive gene pool. Notwithstanding these results, new taxa still emerge.

A. papilionacea ssp papilionacea was first described by Bateman, Pridgeon and Chase in 1997 and was originally considered to have an extensive distribution through most of central and southern Europe. The burgeoning number of newly described taxons has however diminished this range considerably and the most recent of these reclassifications
has had the greatest affect. In 2017, Kreutz described A. papilionacea v septentrionalis, this being the nomenclature for a taxon from the more northerly regions of A. papilionacea's former range. This new sub-species is found in south east France, northern Italy, the Balkans, northern Greece, Bulgaria and Romania. As far as can be determined therefore, this leaves A. papilionacea ssp papilionacea distributed in a southerly band from the Iberian peninsula and north Africa, through southern Italy and across the Adriatic to the Peloponnese.

When mature, A. papilionacea ssp papilionacea is a lax flowered orchid rarely carrying more than 12 medium sized flowers with a pale pink, scalloped lip over layed by dark pink, fan like striations. These markings are usually in broken lines but can sometimes take the form of streaks, dots or rarely be completely absent. The lateral lip margins are gently reflexed forward and the hood is normally deep pink with darker lining, invariably forward pointing and often obscuring the stigmatic cavity.

A. papilionacea ssp papilionacea enjoys a long flowering period which can commence as early as mid January and still be found in May at higher altitudes and in more temperate climes. It favours a position in full sun, on alkaline soils but will thrive on either dry or moist substrates. It is however a species easily crowded out by competing, rank vegetation and prefers a habitat comprised of poor grassland or bare, rocky garrigue.

The assumed distribution of this taxon has already been outlined and its range undoubtedly brings it into contact with other closely related members of its group, notably A. papilionacea  ssp grandiflora (expansa) and A. papilionacea ssp papilionacea f vexillifera. Many authorities do not accept any distinction between these two taxons, sharing as they do,  a broad shouldered, very wide lip which is approximately twice the width of A. papilionacea ssp papilionacea.

Recognizing the various sub-species, varieties and forms of this taxon is difficult in the extreme and many  argue with some cogencey that these differences simply reflect its polymorphic development. Although some of the distinctions are tenuous and explicable via natural variation, others are morphologically at least, more profound and perhaps represent justifiable separation.






















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